BACTERIAL PRODUCTION (THYMIDINE UPTAKE) IN MCMURDO DRY VALLEY LAKES


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BACTERIAL PRODUCTION (THYMIDINE UPTAKE) IN MCMURDO DRY VALLEY LAKES
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PRINCIPAL INVESTIGATOR: John Priscu
Address:Department of Biology, Montana State University, Bozeman MT, 59717
Phone:(406)994-3250
E-Mail:ubijp@gemini.oscs.montana.edu or JPriscu@LTERnet.edu

KEYWORDS: lake, limnology, thymidine, bacteria, production, Antarctica

ABSTRACT: An important part of the McMurdo Long Term Ecological Research (LTER) is monitoring of spatial and temporal patterns, and processes that control bacterial production in perennial ice-covered lakes. This data set quantifies thymidine uptake by bacteria and can be used to estimate bacterial production.

VARIABLES: limno run, location name, location code, date, depth (m), TDR (nM TDR/day), TDR comments, file name

RESEARCH LOCATION: Samples were collected from the East Lake Bonney, West Lake Bonney, Lake Hoare, and Lake Fryxell limnological sampling stations, located in the McMurdo Dry Valleys of Antarctica. Descriptions of these lakes can be found in the McMurdo Dry Valley Lake Descriptions file.

METHODS: Lake water samples were collected at specific depths with a five-liter Niskin bottle during normal LTER limnological sampling. Sub-samples were decanted into three 1-L Nalgene bottles (2-light and 1-amber), two-500 mL amber Nalgene bottles, three-150 mL borosilicate glass bottles, two-20 mL scintillation vials, and one-30 mL serum vial. Five-20 mL scintillation vials (3-live treatments, 2-kill treatments) were prepared at each depth for TDR analysis. TDR samples (10 mL) were taken from the one-liter amber Nalgene bottle and placed in each scintillation vial. 0.5 mL filtered formalin (0.2µm) was pipetted into each killed treatment vial. 3H Thymidine was pipetted into each vial (final concentration of 20 nM thymidine), first the live treatments and then the kill treatments. Samples were incubated in the dark at 1-4°C for 20 hours (thymidine incorporation has been shown to be linear for 20 hours under these conditions, Priscu unpublished). At the end of the incubation period 10 mL of ice cold 10% TCA was added to stop thymidine uptake and precipitate nucleotides. Vials were stored at 4°C until filtered. Each sample was filtered through a 0.2 µm Nucleopore polycarbonate membrane filter. Both the scintillation vial and filter tower were rinsed three times with 5% TCA. Each filter was placed into a new scintillation vial and 15 mL of Cytoscint cocktail was added. Samples were counted with a pre-calibrated (3H) liquid scintillation counter. Thymidine uptake rate (TDR nM d-1) was calculated using the following equation:

TDR = (DPML - DPMK) * a
alpha * t

where DPML is the average dpm for the live treatment, DPMK is the average dpm for the kill treatment, a is the final concentration of thymidine in each vial (20nM), alpha is the total dpm added to each vial, and t is the incubation period. The thymidine uptake rate was adjusted to the ambient lake temperature using the Arrhenius equation:

TDRadj = TDR * e (Ea (( 1 / (CI + 273 K)) - ( 1 / (CA + 273 K))) / R )

where Ea is the energy of activation (12,600 cal mol-1, Q10 = 2.2), °CI is the incubation temperature (°C), °CA is the ambient lake water temperature at specific depth, and R is a gas constant (1.987 cal mol-1 K-1).

TIMING: Samples were collected from the following sites and dates:

 
Season		East Bonney	West Bonney	Hoare		Fryxell
1993-1994	27-Oct-1993	29-Oct-1993	 6-Nov-1993	16-Nov-1993
 		10-Nov-1993	12-Nov-1993
 		24-Nov-1993	26-Nov-1993	 1-Dec-1993	 7-Dec-1993
 		 7-Dec-1993	 9-Dec-1993
 		21-Dec-1993	23-Dec-1993	30-Dec-1993	 7-Jan-1994
 													
1994-1995	 2-Nov-1994	 5-Nov-1994	10-Nov-1994	14-Nov-1994
 		23-Nov-1994	25-Nov-1994	30-Nov-1994	 4-Dec-1994
 		17-Dec-1994	19-Dec-1994	24-Dec-1994	30-Dec-1994
 		10-Jan-1995	12-Jan-1995	20-Jan-1995	17-Jan-1995
 		
1995-1996	19-Sep-1995	21-Sep-1995	 7-Sep-1995	14-Sep-1995
 						14-Sep-1995
 		 6-Oct-1995	 4-Oct-1995	25-Sep-1995	28-Sep-1995
 		17-Oct-1995	19-Oct-1995	10-Oct-1995	14-Oct-1995
 		 2-Dec-1995	 5-Dec-1995	21-Nov-1995	26-Nov-1995
 		 1-Jan-1996	 4-Jan-1996	23-Dec-1995	27-Dec-1995
 		
1996-1997	 3-Nov-1996	 1-Nov-1996	 8-Nov-1996	26-Oct-1996
 		24-Nov-1996	26-Nov-1996	 2-Dec-1996	19-Nov-1996
 		15-Jan-1997	17-Jan-1997	12-Jan-1997	 9-Jan-1997

1997-1998	 8-Nov-1997	11-Nov-1997	15-Nov-1997	18-Nov-1997
 		20-Dec-1997	22-Dec-1997	26-Dec-1997	29-Dec-1997

1998-1999	 9-Nov-1998	11-Nov-1998	 3-Nov-1998	28-Oct-1998
		 6-Dec-1998	 9-Dec-1998 	 1-Dec-1998	23-Nov-1998
		30-Dec-1998	 1-Jan-1999	26-Dec-1998	20-Dec-1998

1999-2000	 6-Nov-1999	11-Nov-1999	17-Nov-1999	30-Oct-1999
		 3-Dec-1999	 5-Dec-1999	12-Dec-1999	27-Nov-1999
		28-Dec-1999	30-Dec-1999	 4-Jan-2000	23-Dec-1999

CITATIONS:

COMMENTS: The energy of activation was experimentally derived for Lake Bonney (Priscu, unpublished). If the average of the kill treatment uptake exceeds live treatment uptake, uptake is reported as zero.

STATUS: Public Access (Type 1).

VARIABLE DESCRIPTION:

VARIABLE

TYPE

DESCRIPTION

UNITS

MISSING VALUE INDICATOR

MINIMUM

MAXIMUM

PRECISION

Limno Run

Text

Code for lake's sampling location and date

none

Required entry

n/a

n/a

n/a

Location Name

Text

Name of lake where measurement was made

none

Required entry

n/a

n/a

n/a

Location Code

Text

Code for site where measurement was made

none

Required entry

n/a

n/a

n/a

Date

Date

Date on which sample was gathered

mm/dd/yy

Required entry

01/01/93

12/31/99

dd

Depth (m)

Number

Distance below ice from which sample was drawn

meters

Required entry

1

25

1

TDR (nM TDR/d)

Number

Thymidine uptake rate using tritiated thymidine

nM TDR/day

Null

0

1

0.001

TDR Comments

Text

Helpful hints about the sample

none

Null

n/a

n/a

n/a

File Name

Text

Name of file in which data was submitted

none

Null

n/a

n/a

n/a

LOG: Data from this table was submitted to INSTAAR by John Priscu's team at Montana State University. The raw data files listed under 'file name' are the names of the original files submitted. The 1993/94 and 1994/95 datasets are Microsoft Excel version 6.0 files, and the 1995/96, 1996/97 and 1997/98 datasets are ascii text files.

Upon arrival at INSTAAR, the data manager fine-tuned the location codes and limno runs to match those provided in the "locations, dates, codes for lake chemistry, biology samples" file. The file was imported into Microsoft Access on INSTAAR's Unix system, and can currently be found there. The file was then exported in ascii, comma delimited text and MS-DOS text (table layout) on the MCM LTER web site. Both of these files are linked to this web page above.

Information for the metadata was obtained from the Metatdr9697.rtf file. The file was called up using Microsoft Word version 6.0. Text from this file was used to create this page in html format.

NOTE: Data contained in these files has been subjected to quality control standards imposed by the investigator. The user of this data should be aware that, while efforts have been taken to ensure that these data are of the highest quality, there is no guarantee of perfection for the data contained herein and the possibility of errors exists. If you encounter questionable data, please contact the MCM LTER data manager (; (303)492-4639) so that the data can be corrected or qualified. Thus, these data may be modified and future data will be appended.